a binary presence-absence matrix in which rows are species and columns
are sites. The entries may be either abundances or occurrences of indivdual species.
Value
Returns the variance ratio of the matrix.
Details
The variance ratio is the ratio of the variance in species number
among sites to the sum of the variance of the species occurrences. If the average
covariation in abundance (or occurrence) of each species pair is close to zero,
the expected value for this ratio is approximately 1.0. V-ratios larger than 1.0
imply positive average covariation in the abundance of species pairs, whereas V-ratios
significantly smaller than 1.0 imply negative average covariation.
References
Schluter, D. 1984. A variance test for detecting species associations,
with some example applications. Ecology 65: 998-1005.
McCulloch, C.E. 1985. Variance tests for species association. Ecology 66: 1676-1681.