Calculates Pielou's measure of species evenness, i.e. J = H'/ln(S) where H' is
Shannon Weiner diversity and S is the total number of species in a sample, across all samples in dataset.
Usage
evenness(x)
Arguments
x
A vector or matrix of species abundances (e.g. counts). The
function assumes that species are in columns and sites are in rows.
Value
Returns Pielou's J.
Details
Many diveristy indices incoporate evenness (e.g. Simpson's diversity, Shannon-
Weiner diversity). Diversity indices which concentrate totally on evenness are
fraught with problems including dependence on species counts (McCune and Grace
2002). A particular problem with Pielou's index is that it is a ratio of a
relatively stable index, H', and one that is strongly dependent on sample size, S.
References
McCune, B., and Grace, J.B. (2002) Analysis of ecological communities. MjM Software
design. Gelenden Beach OR.