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phytools (version 1.0-3)

evol.vcv: Likelihood test for variation in the evolutionary variance-covariance matrix

Description

This function takes an object of class "simmap" with a mapped binary or multistate trait and data for an arbitrary number of continuously valued character. It then fits the multiple evolutionary variance-covariance matrix (rate matrix) model of Revell & Collar (2009; Evolution).

Usage

evol.vcv(tree, X, maxit=2000, vars=FALSE, ...)

Value

An object of class "evol.vcv" with the following components:

R.single

vcv matrix for the single rate matrix model.

vars.single

optionally, a matrix containing the variances of the elements of R.single.

logL1

log-likelihood for single matrix model.

k1

number of parameters in the single marix model.

R.multiple

m x m x p array containing the p estimated vcv matrices for the p regimes painted on the tree.

vars.multiple

optionally, an array containing the variances of the parameter estimates in R.multiple.

logL.multiple

log-likelihood of the multi-matrix model.

k2

number of parameters estimated in this model.

P.chisq

P-value of the \(\chi^2\) test on the likelihood ratio.

convergence

logical value indicating whether or not the optimization has converged.

Arguments

tree

an object of class "simmap". If tree is an object of class "phylo" then a simple multivariate Brownian motion model will be fit to the data in X.

X

an n x m matrix of tip values for m continuously valued traits in n species - row names should be species names. If X is supplied as a data frame it will be coerced into a matrix without warning.

maxit

an optional integer value indicating the maximum number of iterations for optimization. This quantity may need to be increased for difficult optimizations.

vars

an optional logical value indicating whether or not to estimate the variances of the parameter estimates from the Hessian matrix.

...

optional arguments. The most important optional argument at this time is error_vcv which should contain a list of matrices of sampling variances and covariances for (and between) the means of each species. The sampling variance for the mean is just the square of the sampling error. Sampling covariances will tend to be zero (or close to zero) if error for different traits is uncorrelated, for instance, because different specimens were used to estimate the means for different traits, and non-zero otherwise.

Author

Liam Revell liam.revell@umb.edu

Details

This function performs optimization by maximizing the likelihood with respect to the Cholesky matrices using optim. Optimization is by method="Nelder-Mead". Using box constraints does not make sense here as they would be applied to the Cholesky matrix rather than the target parameters. Users may have to increase maxit for large trees and/or more than two traits.

References

Revell, L. J., and D. C. Collar (2009) Phylogenetic analysis of the evolutionary correlation using likelihood. Evolution, 63, 1090-1100.

Revell, L. J. (2012) phytools: An R package for phylogenetic comparative biology (and other things). Methods Ecol. Evol., 3, 217-223.

See Also

evol.rate.mcmc, brownie.lite

Examples

Run this code
## load data from Revell & Collar (2009)
data(sunfish.tree)
data(sunfish.data)
## fit multi-correlation model
sunfish.fit<-evol.vcv(sunfish.tree,sunfish.data[,2:3])
print(sunfish.fit)

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