pez (version 1.0-0)

dispersion: Calculate (phylogenetic) dispersion: examine assemblages in the context of a source pools

Description

As described in Pearse et al. (2014), an evenness metric is one the examines the phylogenetic structure of species present in each assemblage in the context of a source pool of potentially present species. Unlike other metrics, the values of a dispersion metric is *contingent* on the definition of source pool, and (often) randomisations used to conduct that comparison. For completeness, options are provided to calculate these metrics using species traits.

Usage

dispersion(data, metric = c("all", "sesmpd", "sesmntd", "sespd", "innd", "d"),
  permute = 1000, null.model = c("taxa.labels", "richness", "frequency",
  "sample.pool", "phylogeny.pool", "independentswap", "trialswap"),
  abundance = FALSE, sqrt.phy = FALSE, traitgram = NULL,
  traitgram.p = 2, ext.dist = NULL, ...)

Arguments

data

comparative.comm object

permute

number of null permutations to perform (default 1000)

metric

default (all) calculates everything; individually call-able metrics are: SESmpd, SESmntd, SESpd, innd, d.

null.model

one of "taxa.labels", "richness", "frequency", "sample.pool", "phylogeny.pool", "independentswap", or "independentswap". These correspond to the null models available in picante; only d does no

abundance

Whether to use abundance-weighted forms of these metrics (default: FALSE). D, which is presence/absence only, and so will not be calculated when TRUE.

sqrt.phy

If TRUE (default is FALSE) your phylogenetic distance matrix will be square-rooted; specifying TRUE will force the square-root transformation on phylogenetic distance matrices (in the spirit of Leitten and Cornwell, 2014). See `details' for details about

traitgram

If not NULL (default), a number to be passed to funct.phylo.dist (phyloWeight; the `a' parameter), causing analysis on a distance matrix reflecting both traits and phylogeny (0 --> only phylogeny, 1 --> only traits; see fun

ext.dist

Supply an external species-level distance matrix for use in calculations. See `details' for comments on the use of distance matrices in different metric calculations.

traitgram.p

A value for `p' to be used in conjunction with traitgram when calling funct.phylo.dist.

...

additional parameters to be passed to metrics (unlikely you will want to use this!)

Value

• a phy.structure list object of metric values

Details

Most of these metrics do not involve comparison with some kind of evolutionary-derived expectation for phylogenetic shape. Those that do, however, such as D, make no sense unless applied to a phylogenetic distance matrix - their null expectation *requires* it. Using square-rooted distance matrices, or distance matrices that incorporate trait information, can be an excellent thing to do, but (for the above reasons), pez won't give you an answer for metrics for which WDP thinks it makes no sense. SESpd can (...up to you whether it should!...) be used with a square-rooted distance matrix, but the results *will always be wrong* if you do not have an ultrametric tree (branch lengths proportional to time) and you will be warned about this. WDP strongly feels you should only be using ultrametric phylogenies in any case, but code to fix this bug is welcome.

References

Pearse W.D., Purvis A., Cavender-Bares J. & Helmus M.R. (2014). Metrics and Models of Community Phylogenetics. In: Modern Phylogenetic Comparative Methods and Their Application in Evolutionary Biology. Springer Berlin Heidelberg, pp. 451-464.

sesmpd,sesmntd Webb C.O. (2000). Exploring the phylogenetic structure of ecological communities: An example for rain forest trees. American Naturalist, 156, 145-155.

sespd Webb C.O., Ackerly D.D. & Kembel S.W. (2008). Phylocom: software for the analysis of phylogenetic community structure and trait evolution. Bioinformatics Applications Note, 24, 2098-2100.

innd Ness J.H., Rollinson E.J. & Whitney K.D. (2011). Phylogenetic distance can predict susceptibility to attack by natural enemies. Oikos, 120, 1327-1334.

d Fritz S.A. & Purvis A. (2010). Selectivity in Mammalian Extinction Risk and Threat Types: a New Measure of Phylogenetic Signal Strength in Binary Traits. Conservation Biology, 24, 1042-1051.

See Also

shape evenness dissimilarity

Examples

data(laja)
data <- comparative.comm(invert.tree, river.sites, invert.traits)
dispersion(data)
dispersion(data, metric = "sesmpd", permute = 100)